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TONGIORGI E. - 1935. Un primo contributo alla cariologia delle Inuleae. Rendiconto delle sedute e brevi comunicazioni.. N. Giorn. Bot. Ital., n.s. , 42: 261-262.
Magulaev, A. Y. 1982. The number of chromosomes of the species of Asteraceae, Caryophyllaceae and Plantaginaceae of the North Caucasus. Biol. Nauki (Moscow) 11 (227): 74–79.